The game begins in the year 2017, with U.S. Army Captain Elias Walker (Stephen Lang) telling his two sons, Logan and David "Hesh" Walker (Brandon Routh) the legend of how the Ghosts, an elite coalition of all US Special Operations units, first appeared. Meanwhile, in space, Federation soldiers hijack the controlling space station of the Orbital Defense Initiative (ODIN), an orbital superweapon that utilizes kinetic bombardment, and use it to destroy several cities in the Southwestern United States. The surviving American astronauts, Baker and Mosley, sacrifice themselves to self-destruct the space station and prevent the other ODIN satellites from launching their payloads further inland. Elias and the teenage Logan and David narrowly escape the destruction of San Diego.
Ten years later, the United States and the Federation fight to a bloody stalemate along a front that comprises the destroyed cities, also known as "No Man's Land", but it has become a war of attrition as more Federation invasion forces attempt to break the frontlines. However, the Americans stand their ground, fighting near the former US-Mexico border. Logan and Hesh are part of a U.S special unit under the command of Elias and are accompanied by their loyal military-trained German Shepherd, Riley. During a patrol, they spot an American working with the Federation, named Gabriel Rorke (Kevin Gage). Soon after, the brothers are ambushed by wolves, but are rescued by Thomas Merrick and Keegan Russ, members of the Ghosts, who are aiming to rescue their companion Ajax, who was captured by Rorke. Logan and Hesh join the mission to rescue Ajax, but arrive too late to rescue him as he is killed by the Federation.
The brothers return to Santa Monica, California, where the Federation forces invade the city as U.S forces holdout against their assault. As the Americans counterattack, Hesh and Logan reunite with their father, who reveals himself as the leader of Ghosts. Knowing that his sons are qualified to join his special unit, Elias recruits them into the Ghosts, and they learn from him that Rorke was once their leader and Elias' best friend. However, during a successful mission to assassinate General Diego Almagro, then president of the Federation in the capital city Caracas, Elias is forced to abandon Rorke, who was presumed dead, but instead turned traitor after being captured and brainwashed by the Federation and began hunting down members of the Ghosts.
The Ghosts track down and capture Rorke, but while flying back home, Rorke's men attack their plane and rescue him. The team then is forced to land deep into the Amazon Jungle, where they witness a rocket being launched for an unknown purpose. Once reunited and rescued, the Ghosts raid a Federation laboratory in the Andes and obtain data regarding classified information in a factory at Rio De Janeiro. To secure a path to the factory, the Ghosts destroy the Federation's Atlas oil platform located in Antarctica to lure the enemy fleet away from their objective and sink a destroyer guarding the Brazilian coast. Once inside the factory, the team discovers that the Federation had scuffled and reverse engineered ODIN's technology and developed their own orbital bombardment system code named "LOKI". After destroying the factory, Elias and his sons regroup at a safehouse in Las Vegas, but are captured by Rorke. After Rorke kills Elias, the brothers manage to escape along with Merrick, Keegan and Riley.
Once the Federation's LOKI becomes operational, the United States's defeat will become inevitable. In response, the United States decides to pool all their remaining forces, including the last aircraft carrier, the USS Liberator, in a synchronized all-out assault to take down an enemy space center in Chile, while a small team of soldiers boards a shuttle to take over the Federation satellites in space. Once both objectives are completed, Hesh and Logan pursue Rorke to avenge their father and are seemingly successful, only to find out during the aftermath that Rorke survived. Logan attempts to defend his wounded brother, but has his arm broken by Rorke, who announces his plans to brainwash Logan into killing the rest of the Ghosts before kidnapping Logan.
In a post-credits scene, Logan is seen being kept inside a pit in the jungle, presumably going through the same torture methods that Rorke went through.
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